PDFs disponibles sur demande par mail : leroy.boris at

International Peer-Reviewed Journals


49. Ahmed DA, Hudgins EJ, Cuthbert RN, Kourantidou M, Diagne C, Haubrock PJ, Leung B, Liu C, Leroy B, Petrovskii S, Courchamp F. Managing biological invasions: the cost of inaction. Preprint available 10.21203/

48. Morel L, Jung V, Leroy B, Villéger S, Clément B, Schnitzler A, Ysnel F. Changes in community assembly processes during succession increase biodiversity and affect plant community trajectories. Under consideration

47. Leroy B, Kramer AV, Vaissière AC, Courchamp F, Diagne C. Analysing global economic costs of invasive alien species with the invacost R package. Preprint available 10.1101/2020.12.10.419432


46. Robuchon M, Pavoine S, Delli G, Faith DP, Mandrici A, Pellens R, Véron S, Dubois G, Leroy B. 2021. Revisiting species and areas of interest for conserving global mammalian phylogenetic diversity. Nature Communications 12:3694 10.1038/s41467-021-23861-y

45. Bradshaw CJA, Hoskins AJ, Haubrock PJ, Cuthbert RN, Diagne C, Leroy B, Andrews L, Page B, Cassey P, Sheppard AW, Courchamp F. 2021. Detailed assessment of the reported economic costs of invasive species in Australia. In: Zenni RD, McDermott S, García-Berthou E, Essl F (Eds) The Economic costs of biological invasions in the world. NeoBiota 67: 511–550. 10.3897/neobiota.67.58834

44. Bourdaud P, Ben Rais Lasram F, Araignous E, Champagnat J, Grusd S, Halouani G, Hattab T, Leroy B, Noguès Q, Raoux A, Safi G, Niquil N. 2021. Impacts of climate change on the Bay of Seine ecosystem: forcing a spatio-temporal trophic model with predictions from an ecological niche model. Fisheries Oceanography 10.1111/fog.12531

43. Crystal-Ornelas R, Hudgins EJ, Cuthbert RN, Haubrock PJ, Fantle-Lepczyk J, Angulo E, Kramer AM, Ballesteros-Mejia L, Leroy B, Leung B, López-López E, Diagne C, Courchamp F. 2021. Economic costs of biological invasions within North America. In: Zenni RD, McDermott S, García-Berthou E, Essl F (Eds) The Economic costs of biological invasions in the world. NeoBiota 67: 485–510. 10.3897/neobiota.67.58038

42. Diagne C, Turbelin AJ, Moodley D, Novoa A, Leroy B, Angulo E, Adamjy T, Dia CAKM, Taheri A, Tambo J, Dobigny G, Courchamp F. 2021. The economic costs of biological invasions in Africa: a growing but neglected threat? In: Zenni RD, McDermott S, García-Berthou E, Essl F (Eds) The Economic costs of biological invasions in the world. NeoBiota 67: 11–51. 10.3897/neobiota.67.59132

41. Renault D, Manfrini E, Leroy B, Diagne C, Ballesteros-Mejia L, Angulo E, Courchamp F (2021) Biological invasions in France: Alarming costs and even more alarming knowledge gaps. In: Zenni RD, McDermott S, García-Berthou E, Essl F (Eds) The Economic costs of biological invasions in the world. NeoBiota 67: 191–224. 10.3897/neobiota.67.59134

40. Kourantidou M, Cuthbert RN, Haubrock PJ, Novoa A, Taylor NG, Leroy B, Capinha C, Renault D, Angulo E, Diagne C, Courchamp F. 2021. Economic costs of invasive alien species in the Mediterranean basin. In: Zenni RD, McDermott S, García-Berthou E, Essl F (Eds) The Economic costs of biological invasions in the world. NeoBiota 67: 427–458. 10.3897/neobiota.67.58926

39. Haubrock PJ, Turbelin AJ, Cuthbert RN, Novoa A, Taylor NG, Angulo E, Ballesteros-Mejia L, Bodey TW, Capinha C, Diagne C, Essl F, Golivets M, Kirichenko N, Kourantidou M, Leroy B, Renault D, Verbrugge L, Courchamp F. 2021. Economic costs of invasive alien species across Europe In: Zenni RD, McDermott S, García-Berthou E, Essl F (Eds) The Economic costs of biological invasions in the world. NeoBiota 67: 153–190. 10.3897/neobiota.67.58196

38. Nogues Q, Raoux A, Araignous E, Chaalali A, Hattab T, Leroy B, Ben Rais Lasram F, David V, Le Loc’h F, Dauvin JC, Niquil N. 2021. Cumulative Effects of Marine Renewable Energy and Climate Change on Ecosystem Properties: Sensitivity of Ecological Network Analysis. Ecological Indicators IN PRESS 10.1016/j.ecolind.2020.107128

37. Diagne C, Leroy B, Vaissière AC, Gozlan RE, Roiz D, Jarić I, Salles JM, Bradshaw CJA & Courchamp F. 2021. High and rising economic costs of biological invasions worldwide Nature 592, pages 571–576 10.1038/s41586-021-03405-6


36. Schickele A, Goberville E, Leroy B, Beaugrand G, Hattab T, Francour P & Raybaud V. 2020. European small pelagic fish distribution under global change scenarios. Fish and Fisheries 22:212-225 10.1111/faf.12515

35. Godet C, Robuchon M, Leroy B, Cotté C, Baudena A, Da Silva O, Fabri-Ruiz S, Lo Monaco C, Sergi S, Koubbi P. 2020. Matching zooplankton abundance and environment in the South Indian Ocean and Southern Ocean. Deep Sea Research Part I: Oceanographic Research Papers 163, 103347 10.1016/j.dsr.2020.103347

34. Diagne C, Leroy B, Gozlan R, Vaissière AC, Assailly C, Nuninger L, Roiz D, Jourdain F, Jaric I, Courchamp F. 2020. INVACOST, a public database of the economic costs of biological invasions worldwide. Nature Scientific Data 7, 277 10.1038/s41597-020-00586-z

33. Ben Rais Lasram F, Hattab T, Nogues Q, Beaugrand G, Dauvin, JC, Halouani, G, Le Loc’H F, Niquil N, Leroy B. 2020. An open-source framework to model present and future marine species distributions at local scale. Ecological Informatics 59:101130 10.1016/j.ecoinf.2020.101130

32. Louppe V, Leroy B, Herrel V, Veron G. 2020. The globally invasive small Indian mongoose Urva auropunctata is likely to spread with climate change. Scientific Reports 10:7461 10.1038/s41598-020-64502-6

31. Schickele A, Leroy B, Beaugrand G, Goberville E, Hattab T, Francour P, Raybaud V. 2020. Modelling European small pelagic fish distribution: Methodological insights. Ecological Modelling 416:108902 10.1016/j.ecolmodel.2019.108902


30. Leroy B, Dias MS, Giraud E, Hugueny B, Jézéquel C, Leprieur F, Oberdorff T, Tedesco PA. 2019. Global biogeographical regions of freshwater fish species. Journal of Biogeography, 46:2407-2419 10.1111/jbi.13674

29. Louppe V, Leroy B, Herrel V, Veron G. 2019. Current and future climatic regions favourable for a globally introduced wild carnivore, the raccoon Procyon lotor. Scientific Reports 9:9174 10.1038/s41598-019-45713-y

28. Meynard CN, Kaplan DM, Leroy B. Testing methods in species distribution modelling using virtual species: what have we learnt and what are we missing? Ecography Special Issue: E4 award 42:2021-2036 10.1111/ecog.04385

27. Meynard CN, Kaplan DM, Leroy B. 2019. Detecting outliers in species distribution data: some caveats and clarifications on a virtual species study. Journal of Biogeography 46:2141-2144 10.1111/jbi.13626

26. Robuchon M, Faith DP, Julliard R, Leroy B, Pellens R, Robert A, Thévenin C, Veron S, Pavoine S. 2019. Species splitting increases estimates of evolutionary history at risk. Biological Conservation 235:27-35 10.1016/j.biocon.2019.03.041

25. Robuchon M, Leroy B, Jézéquel C, Hugueny B. 2019. Correlations between broad-scale taxonomic and genetic differentiations suggest a dominant imprint of historical processes on beta diversities. Journal of Biogeography 46:1083-1095 10.1111/jbi.13559

24. Morel L, Dujol B, Courtial C, Vasseur M, Leroy B, Ysnel F. 2019. Spontaneous recovery of functional diversity and rarity of ground-living spiders shed light on the conservation importance of recent woodlands. Biodiversity and Conservation 28:687–709 10.1007/s10531-018-01687-3


23. Leroy B, Delsol R, Meynard CN, Hugueny B & Bellard C. 2018. Without quality presence-absence data, discrimination metrics such as TSS can be misleading measures of model performance. Journal of Biogeography, 45:1994-2002 10.1111/jbi.13402

22. Bellard C, Jeschke J, Leroy B & Mace G.M. 2018. Insights from modelling studies on how climate change affects invasive alien species geography. Ecology & Evolution 8:5688-5700 10.1002/ece3.4098

21. Pétillon J, Leroy B, Djoudi EA & Vedel V. 2018. Small and large spatial scale coexistence of Ctenid spiders in a neotropical forest (French Guiana). Tropical Zoology. 31:85-98 10.1080/03946975.2018.1448531

20. Mammola S & Leroy B. 2018. Applying species distribution models to caves and other subterranean ecosystems. Ecography 41:1194-1208 10.1111/ecog.03464

19. Hervé, V, Leroy B, Da Silva Pires A & Lopez, PJ. 2018. Aquatic urban ecology at the scale of a capital: community structure and interactions in street gutters. The ISME Journal 12:253-266 10.1038/ismej.2017.166


18. Bellard C, Rysman J-F, Leroy B, Claud C & Mace GM. 2017. A global picture of biological invasion threat on islands. Nature Ecology & Evolution 1:1862-1869 s41559-017-0365-6

17. Gillard M, Thiebaut G, Deleu C & Leroy B. 2017. Present and future distribution of three aquatic plant taxa across the world: decrease in native and increase in invasive ranges. Biological Invasions 19:2159-2170. 10.1007/s10530-017-1428-y

16. Hawkins BA, Leroy B, Rodríguez MÁ, Singer A, Villalobos F, Wang X, & Zelený D. 2017. Structural bias in aggregated trait variables driven by repeated species co-occurrences: a pervasive problem in community and assemblage data. Journal of Biogeography 44:1199-1211. 10.1111/jbi.12953

15. Leroy B, Gallon R, Feunteun E, Robuchon M & Ysnel F. 2017. Cross-taxon congruence in the rarity of subtidal rocky marine assemblages : no taxonomic shortcut for conservation monitoring. Ecological Indicators 77:239-249 10.1016/j.ecolind.2017.02.012


14. Bradshaw CJA, Leroy B, Bellard C, Albert C, Roiz D, Barbet-Massin M, Fournier A, Salles JM, Simard F & Courchamp F. 2016. Massive yet grossly underestimated global costs of invasive insects. Nature Communications 7:12986 10.1038/ncomms12986

13. Bellard C, Leroy B, Thuiller W, Rysman JF & Courchamp F. 2016. Major drivers of invasions risks throughout the world. Ecosphere 7:e01241 10.1002/ecs2.1241

12. Leroy B, Meynard CN, Bellard C, Courchamp F. 2016. virtualspecies: an R package to generate virtual species distributions. Ecography 39:599-607 10.1111/ecog.01388


11. Potier S, Carpentier A, Grémillet D, Leroy B, Lescroël, A. 2015. Individual repeatability of foraging behaviour in a generalist marine predator. Animal Behaviour 103:83-90 10.1016/j.anbehav.2015.02.008


10. Gallon R, Robuchon M, Leroy B, Le Gall L, Valero M, Feunteun E. 2014. 20 years of observed and predicted changes in subtidal red seaweed assemblages along a biogeographical transition zone: inferring potential causes from environmental data. Journal of Biogeography 41:2293-2306 10.1111/jbi.12380

9. Bellard C, Leclerc C, Leroy B, Bakkenes M, Veloz S, Thuiller W & Courchamp F. 2014. Vulnerability of biodiversity hotspots to global change. Global Ecology and Biogeography 23:1376–1386 10.1111/geb.12228

8. Leroy B, Le Viol I, Pétillon J. 2014. Complementarity of rarity, specialisation and functional diversity metrics to assess community responses to environmental changes, using an example of spiders in salt marshes. Ecological Indicators 46:351-357 j.ecolind.2014.06.037

7. Puzin C, Leroy B & Pétillon J. 2014. Intra- and inter-specific variation in size and habitus of two sibling spider species (Araneae: Lycosidae): taxonomic and biogeographic insights from sampling across Europe. Biological Journal of the Linnean Society 113:85-96 10.1111/bij.12303

6. Leroy B, Bellard C, Dubos N, Colliot A, Vasseur M, Courtial C, Bakkenes M, Canard A & Ysnel F. 2014. Forecasted climate and land use changes, and protected areas: the contrasting case of spiders. Diversity and Distributions 20:686-697 10.1111/ddi.12191


5. Bellard C, Thuiller W, Leroy B, Genovesi P, Bakkenes M & Courchamp F. 2013. Will climate change promote future invasions? Global Change Biology 19:3740-3748 10.1111/gcb.12344

4. Leroy B, Canard A & Ysnel F. 2013. Integrating multiple scales in rarity assessments of invertebrate taxa. Diversity and Distributions 19:794-803 10.1111/ddi.12040

3. Leroy B, Paschetta M, Canard A, Bakkenes M, Isaia M & Ysnel F. 2013. First assessment of effects of environmental change on threatened spiders: potential impacts on Dolomedes plantarius (Clerck) and its conservation plans. Biological Conservation 161:155-163 10.1016/j.biocon.2013.03.022


2. Leroy B, Pétillon J, Gallon R, Canard A & Ysnel F. 2012. Improving occurrence-based rarity metrics in conservation studies by including multiple rarity cut-offs. Insect Conservation and Diversity 5:159-168 10.1111/j.1752-4598.2011.00148.x


1. Leroy B, Morel L, Eybert MC, Ysnel F & Georges A. 2011. Rehabilitation project of a managed marsh: biodiversity assessment of different management measures. Procedia Environmental Sciences 9:96-103 10.1016/j.proenv.2011.11.016

Software / packages

Leroy B. 2020. INVACOST, package for the R software. INVACOST Database With Methods To Analyse Invasion Costs. Current version: 0.1-3 (beta). Release is planned very soon!

Leroy B. 2019-2020. biogeonetworks, package for the R software. Biogeographical networks in R. Current version : 0.1.2.

Leroy B. 2014-2019. virtualspecies, package for the R software. A package to generate virtual species distribution for improvements of modeling techniques. Published on the Comprehensive R Archive Network. Current version : 1.5.1.

Leroy B. 2012-2019. Rarity, package for the R software. A package to calculate rarity indices for species and assemblages of species. Published on the Comprehensive R Archive Network. Current version : 1.3-6.

Expertise reports


Araignous E., Beaugrand G., Ben Rais Lasram F., Bourdaud P., Champagnat J., Dauvin J.-C., Grangeré K., Halouani G., Haraldsson M., Hattab T., Le Loc’h F., Leroy B., Lejart M., Niquil N., Nogues Q., Pagot J.-P., Raoux A., Safi, G. et Villanueva C. Recommandations pour une approche écosystémique des aires d’implantation d’énergies marines renouvelables. Cas d’étude du parc éolien offshore de Courseulles-sur-mer. Plouzané. France Energies Marines Editions, 2019, 60 pages. Edition : Juin 2019


IPBES (2018), The IPBES regional assessment report on biodiversity and ecosystem services for Europe and Central Asia. Rounsevell, M., Fischer, M., Torre-Marin Rando, A. and Mader, A. (eds.). Secretariat of the Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services, Bonn, Germany. 892 pages. 10.5281/zenodo.3237428


Picard L. & Leroy B. 2015. Inventaire des coléoptères aquatiques des mares de Basse-Normandie en vue de l’application d’un indice biologique. Rapport de synthèse 2011-2014. Rapport GRETIA pour le Conseil régional de Basse-Normandie, les Conseils généraux du Calvados, de la Manche et de l’Orne, et l’Agence de l’eau Seine-Normandie. 103 pp. + annexes.


Privet K., Canard A., Courtial C., Djoudi E.A., Leroy B., Vedel, V. & Pétillon J. 2014. Les araignées de la Réserve Naturelle Nationale des Nouragues : Facteurs de variations des assemblages et premier inventaire. Rapport pour la Réserve Naturelle des Nouragues, Guyane Française, 23p

National peer-reviewed journals


1. Pétillon J., Leroy B., Canard A. & Ysnel F. 2010. Les araignées au fil de la gestion. Espaces naturels 31:36-37 [link PDF]